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This is also very true for the air filters on your home furnace/heat pump, if you have one. Even if you aren't running the heat, if you are running the furnace/heat pump's fan to circulate air periodically (which you should be doing to avoid stagnation), air is still passing through the filter. I live in the Seattle area, and our furnace filter comes out absolutely filthy black after every bout of wildfire smoke. It nasty.